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  • Yost, J. M., P. W. Sweeney, E. Gilbert, G. Nelson, R. Guralnick, A. S. Gallinat, E. R. Ellwood, N. Rossington, C.G. Willis et al. 2018. Digitization protocol for scoring reproductive phenology from herbarium specimens of seed plants. Appl. Plant Sciences 6: e1022.
  • Willis, C.G. 2018. REVIEW: The New Ecology: Rethinking a Science for the Anthropocene. By Oswald J. Schmitz. Q. Rev. Biol. 93: 25.
  • Gallinat, A.S., L. Russo, E.K. Melaas, C.G. Willis, and R.B. Primack. 2018. Herbarium specimens show patterns of fruiting phenology in native and invasive plant species across New England. Am. J. Bot. 105: 31-41.
  • Daru, B.H., D.S. Park, R.B. Primack, C.G. Willis, D.S. Barrington, T.J.S. Whitfeld, T.G. Seidler, P.W. Sweeney, D.R. Foster, A.M. Ellison, and C.C. Davis. 2018. Widespread sampling biases in herbaria revealed from large-scale digitization. New Phytol. 217: 939–955.


  • Williams, A., C.G. Willis, C.C. Davis, J. Goh, A.M. Ellison, and E. Law. 2017. Deja Vu: Characterizing worker quality using task consistency. Proceedings of the 5th Conference on Human Computation and Crowdsourcing (HCOMP). Quebec City, Canada.
  • Du, Y., J. Chen, C.G. Willis, Z. Zhou, T. Liu, W. Dai, Y. Zhao, and K. Ma. 2017. Phylogenetic conservatism and trait correlates of spring phenological responses to climate change in northeast China. Ecol. Evol. 7: 6747-6757.
  • Zhang, C., C.G. Willis, J.A. Klein, Z. Ma, J. Li, H. Zhou and X. Zhao. 2017. Recovery of plant species diversity during long-term experimental warming of a species-rich alpine meadow community on the Qinghai-Tibet plateau. Biol. Conserv. 213A: 218-224.
  • Willis, C.G., L. Ellwood, R.B. Primack, C.C. Davis, J.M. Yost, S.J. Mazer, G. Nelson, T.H. Sparks, A. Gallinat, K. Stanley, N.L. Rossington, and P.S. Soltis. 2017. Old plants, new tricks: phenological research using herbarium specimens. Trends Ecol. Evol. 99: 1263-1275
  • Willis, C.G., E. Law, A. Williams, B. Franzone, R. Bernardos, L. Bruno, C. Hopkins, C. Schorn, E. Weber, D. Park, and C.C. Davis. 2017. CrowdCurio: an online crowdsourcing platform to facilitate climate change studies using herbarium specimens. New Phytol. 215: 479-488
  • Rubio de Casas, R., C.G. Willis, W.D. Pearse, C.C. Baskin, J.M. Baskin, and J. Cavender-Bares. 2017. Global biogeography of seed dormancy is determined by seasonality and seed size: a case study in the legumes. New Phytol. 214: 1527-1536
  • Willis, C.G., and K. Donohue. 2017. The evolution of intrinsic reproductive isolation in the genus Cakile (Brassicaceae). J. Evol. Biol. 30: 361–376


  • Davis, C.C., C.G. Willis, B. Connolly, C. Kelly, and A.M. Ellison. 2015. Herbarium records are reliable sources of phenological change driven by climate and provide novel insights into species’ phenological cueing mechanisms. Am. J. Bot. 102: 1599-1609.
  • Willis, C.G. and C.C. Davis. 2015. Rethinking migration. Science. 348: 766.


  • Willis, C.G., B.F. Franzone, Z. Xi, and C.C. Davis. 2014. The establishment of Central American migratory corridors and the biogeographic origins of seasonally dry tropical forests in Mexico. Front. Genet. 5: 433.
  • Willis, C.G., J.C. Hall, R. Rubio de Casas, T.Y. Wang, and K. Donohue. 2014. Diversification and the evolution of dispersal ability in the tribe Brassiceae (Brassicaceae). Ann. Bot. 114: 1675-1686.
  • Zhang, C., C.G. Willis, L.T. Burghardt, W. Qi, K. Liu, P.R. Moura Souza-Filho, Z. Ma, G. Du. 2014. The community‐level effect of light on germination timing in relation to seed mass: a source of regeneration niche differentiation. New Phytol. 204: 496–506.
  • Panchen, Z.A., R.B. Primack, B. Nordt, E.R. Ellwood, A.D. Stevens, S.S. Renner, C.G. Willis, R. Fahey, A. Whittemore, Y. Du, and C.C. Davis. 2014. Leaf out times of temperate woody plants are related to phylogeny, deciduousness, growth habit and wood anatomy. New Phytol. 203: 1208-1219.
  •  Willis, C.G. and C.C. Davis. 2014. Reply to Angelo: Climate change and species loss in Thoreau’s woods (Concord, Massachusetts, USA). Phytoneuron 9: 1-4.
  • Willis, C.G., C.C. Baskin, J.M. Baskin, J.R. Auld, D.L. Venable, J. Cavender-Bares, K. Donohue, R. Rubio de Casas, and The NESCent Germination Working Group. 2014. The evolution of seed dormancy: Environmental cues, evolutionary hubs, and diversification of the seed plants. New Phytol. 203: 300-309.


  •  Willis, C.G. 2013. The role of dispersal and adaptive divergence in the diversification and speciation of the tribe Brassiceae and genus Cakile. Ph.D. dissertation, Duke University.
  • Wolkovich, E.M., T.J. Davies, H. Schaefer, E.E. Cleland, B.I. Cook, S.E. Travers, C.G. Willis, and C.C. Davis. 2013. Temperature-dependent shifts in phenology contribute to the success of exotic species with climate change. Am. J. Bot. 100: 1412-1424.


  • Schneider, B., M. Tobiasz, C.G. Willis, C. Shen. 2012. WALDEN: multi-surface multi-touch simulation of climate change and species loss in Thoreau's woods. Proc. ACM ITS. 387-390.
  • Rubio de Casas, R., K. Donohue, and C.G. Willis. 2012. Plant dispersal phenotypes: a seed perspective of maternal habitat selection. Dispersal Ecology and Evolution. Oxford University Press.


  • Donohue, K., R. Rubio de Casas, L. Burghardt, K. Kovach, and C.G. Willis. 2010. Germination, postgermination adaptation, and species ecological ranges. Ann. Rev. Ecol. Evol. Syst. 41:293-319.
  • Willis, C.G., B.R Ruhfel, B.R. Primack, A.J. Miller-Rushing, J.B. Losos, and C.C. Davis. 2010. Favorable climate change response explains non-native species' success in Thoreau's woods. PLoS One. 5:e8878.
  • Davis, C.C., C.G. Willis, B.R. Primack and A.J. Miller-Rushing. 2010. The importance of phylogeny to the study of phenological response to global climate change. Philos. T. Roy. Soc. B. 365:3201-3213.
  •  Willis, C.G., M.T. Brock, and C. Weinig. 2010. Genetic variation in tolerance of competition and neighbour suppression in Arabidopsis thaliana. J. Evol. Biol. 23:1412-1424.
  •  Willis, C.G., J. Cavender-Bares, M. Halina, C. Lehman, P.B. Reich, A. Keen and S. McCarthy. 2010. Phylogenetic community structure in Minnesota oak savanna is influenced by spatial extent and environmental variation. Ecography. 33:565-577.


  • Willis, C.G., B.R Ruhfel, B.R. Primack, A.J. Miller-Rushing, and C.C. Davis. 2009. Reply to McDonald et al.: Climate change, not deer herbivore, has shaped species decline in Concord, Massachusetts. Proc. Natl. Acad. Sci. 106:E29.


  • Willis, C.G., B.R Ruhfel, B.R. Primack, A.J. Miller-Rushing and C.C. Davis. 2008. Phylogenetic patterns of species loss in Thoreau’s woods are driven by climate change. Proc. Natl. Acad. Sci. 105:17029-17033.


  • Weinig, C., J. Johnston, C.G. Willis and J. N. Maloof. 2007. Antagonistic multilevel selection on size and architecture in variable density settings. Evolution. 61:58–67.